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Fig 1.

Relations between cognitive-, neuronal-, and behavioral states in MCM.

Solid and dotted arrows denote causal and constitutive relations, respectively.

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Fig 2.

State-space diagram with transition probabilities for Example 1.

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Fig 3.

p-values for rejecting the null-hypothesis of Markovianity for each worm (and averaged across worms) as a function of the number of cognitive states.

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Fig 4.

Illustration of the procedure of learning a cognitive state trajectory (see main text for details).

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Fig 5.

Behavioral- and cognitive-behavioral state diagrams of the third worm (see text for details).

Arrows that account for less than 0.1% of outgoing transitions of each node have been removed to reduce clutter.

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Fig 6.

Behavioral- and cognitive-behavioral state diagrams of all worms.

Arrows that account for less than 0.075% of outgoing transitions of each node have been removed to reduce clutter.

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Fig 7.

Side-by-side comparison of the neuronal manifold of the third worm as learned by BunDLe-Net and its representation as a directed graph in the NC-MCM framework.

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Fig 8.

Side-by-side comparison of the neuronal manifold and the cognitive-behavioral state diagram of worm 1.

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Fig 9.

Side-by-side comparison of the neuronal manifold and the cognitive-behavioral state diagram of worm 2.

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Fig 10.

Side-by-side comparison of the neuronal manifold and the cognitive-behavioral state diagram of worm 4.

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Fig 11.

Side-by-side comparison of the neuronal manifold and the cognitive-behavioral state diagram of worm 5.

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Fig 12.

Cognitive-behavioral state transition diagram of the third worm with seven cognitive states.

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Fig 13.

Neuronal perturbations that, when applied during a sustained reversal, the NC-MCM framework predicts to induce a cognitive state transition that increases the probability of maintaining a sustained reversal movement (blue), initiating a ventral turn (orange), or initiating a dorsal turn (green).

Double and single markers indicate the rejection of the null hypothesis of equal means across conditions at significance levels α = 0.01 and α = 0.05, respectively. Plus signs and asterisks indicate significance tests with and without Bonferroni correction, respectively.

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Fig 14.

Neuronal perturbations that, when applied during a slowing movement, the NC-MCM framework predicts to induce a cognitive state transition that increases the probability of maintaining a slowing movement (blue), initiating a forward movement (orange), or initiating a rev2 reversal (green).

Double and single markers indicate the rejection of the null hypothesis of equal means across conditions at significance levels α = 0.01 and α = 0.05, respectively. Plus signs and asterisks indicate significance tests with and without Bonferroni correction, respectively.

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