About the Authors

Shaoke Lou

Contributed equally to this work with: Shaoke Lou, Mingjun Yang

Roles Conceptualization, Formal analysis, Methodology, Visualization, Writing – original draft

Affiliations Program in Computational Biology & Bioinformatics, Yale University, New Haven, Connecticut, United States of America, Department of Molecular Biophysics & Biochemistry, Yale University, New Haven, Connecticut, United States of America

Mingjun Yang

Contributed equally to this work with: Shaoke Lou, Mingjun Yang

Roles Methodology, Visualization

Affiliation School of Electronic Engineering and Computer Science, Queen Mary University of London, Mile End Road, London, United Kingdom

Tianxiao Li

Roles Formal analysis

Affiliations Program in Computational Biology & Bioinformatics, Yale University, New Haven, Connecticut, United States of America, Department of Molecular Biophysics & Biochemistry, Yale University, New Haven, Connecticut, United States of America

Weihao Zhao

Roles Formal analysis

Affiliations Program in Computational Biology & Bioinformatics, Yale University, New Haven, Connecticut, United States of America, Department of Molecular Biophysics & Biochemistry, Yale University, New Haven, Connecticut, United States of America

Hannah Cevasco

Roles Writing – review & editing

Affiliation Program in Computational Biology & Bioinformatics, Yale University, New Haven, Connecticut, United States of America

Yucheng T. Yang

Roles Conceptualization, Supervision

Affiliations Institute of Science and Technology for Brain-Inspired Intelligence, Fudan University, Shanghai, China, MOE Key Laboratory of Computational Neuroscience and Brain-Inspired Intelligence, Fudan University, Shanghai, China, MOE Frontiers Center for Brain Science, Fudan University, Shanghai, China

Mark Gerstein

Roles Project administration, Supervision, Writing – review & editing

mark@gersteinlab.org

Affiliations Program in Computational Biology & Bioinformatics, Yale University, New Haven, Connecticut, United States of America, Department of Molecular Biophysics & Biochemistry, Yale University, New Haven, Connecticut, United States of America, Department of Computer Science, Yale University, New Haven, Connecticut, United States of America, Department of Statistics & Data Science Yale University, New Haven, Connecticut, United States of America, Department of Biomedical Informatics & Data Science, Yale University, New Haven, Connecticut, United States of America

Competing Interests

The authors have declared that no competing interests exist.