About the Authors

Di Zhang

Roles Data curation, Formal analysis, Methodology, Visualization, Writing – original draft

‡ These authors are joint first authors on this work.

Affiliations Center for Quantitative Biology, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing, China, Peking-Tsinghua Center for Life Sciences, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing, China

Sophia Hsin-Jung Li

Roles Conceptualization, Data curation, Resources, Validation

‡ These authors are joint first authors on this work.

Affiliations Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America, Institute of Bioengineering, School of Life Sciences, Swiss Federal Institute of Technology Lausanne, Lausanne, Switzerland, Global Health Institute, School of Life Sciences, Swiss Federal Institute of Technology Lausanne, Lausanne, Switzerland

Christopher G. King

Roles Data curation, Investigation

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Ned S. Wingreen

Roles Conceptualization, Funding acquisition, Investigation, Writing – review & editing

wingreen@princeton.edu (NSW); zgitai@princeton.edu (ZG); zhiyuanli@pku.edu.cn (ZL)

Affiliations Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America, Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

Zemer Gitai

Roles Conceptualization, Funding acquisition, Investigation, Writing – review & editing

wingreen@princeton.edu (NSW); zgitai@princeton.edu (ZG); zhiyuanli@pku.edu.cn (ZL)

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Zhiyuan Li

Roles Conceptualization, Funding acquisition, Project administration, Writing – review & editing

wingreen@princeton.edu (NSW); zgitai@princeton.edu (ZG); zhiyuanli@pku.edu.cn (ZL)

Affiliations Center for Quantitative Biology, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing, China, Peking-Tsinghua Center for Life Sciences, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.