About the Authors

Hongli Ma

Contributed equally to this work with: Hongli Ma, Han Wen

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Software, Writing – original draft, Writing – review & editing

Affiliations Research Center for Mathematics and Interdisciplinary Sciences, Shandong University, Qingdao, China, Terrence Donnelly Centre for Cellular and Biomolecular Research, University of Toronto, Toronto, Ontario, Canada, Department of Molecular Genetics, University of Toronto, Toronto, Ontario, Canada, School of Mathematics, Shandong University, Jinan, China

Han Wen

Contributed equally to this work with: Hongli Ma, Han Wen

Roles Investigation, Methodology, Writing – review & editing

Affiliation Terrence Donnelly Centre for Cellular and Biomolecular Research, University of Toronto, Toronto, Ontario, Canada

Zhiyuan Xue

Roles Software, Visualization

Affiliation West China Biomedical Big Data Center, West China Hospital, Sichuan University, Chengdu, China

Guojun Li

Roles Funding acquisition, Supervision

guojunsdu@gmail.com (GL); Zhaolei.Zhang@utoronto.ca (ZZ)

Affiliations Research Center for Mathematics and Interdisciplinary Sciences, Shandong University, Qingdao, China, School of Mathematics, Shandong University, Jinan, China, School of Mathematical Science, Liaocheng University, Liaocheng, China

Zhaolei Zhang

Roles Conceptualization, Funding acquisition, Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

guojunsdu@gmail.com (GL); Zhaolei.Zhang@utoronto.ca (ZZ)

Affiliations Terrence Donnelly Centre for Cellular and Biomolecular Research, University of Toronto, Toronto, Ontario, Canada, Department of Molecular Genetics, University of Toronto, Toronto, Ontario, Canada, Department of Computer Science, University of Toronto, Toronto, Ontario, Canada

Competing Interests

none