About the Authors

Yu Wang

Roles Formal analysis, Investigation, Methodology, Software, Visualization, Writing – original draft

Affiliations School of Mathematical Sciences, Fudan University, Shanghai, China, Research Institute of Intelligent Complex Systems, Fudan University, Shanghai, China

Fengzhu Sun

Roles Conceptualization, Resources

Affiliation Quantitative and Computational Biology Department, University of Southern California, Los Angeles, California, United States of America

Wei Lin

Roles Formal analysis, Funding acquisition, Methodology, Resources

Affiliations School of Mathematical Sciences, Fudan University, Shanghai, China, Research Institute of Intelligent Complex Systems, Fudan University, Shanghai, China, State Key Laboratory of Medical Neurobiology, MOE Frontiers Center for Brain Science, and Institutes of Brain Science, Fudan University, Shanghai, China, Shanghai Artificial Intelligence Laboratory, Shanghai, China, Key Laboratory of Mathematics for Nonlinear Science (Fudan University), Ministry of Education, Shanghai, China, Shanghai Key Laboratory for Contemporary Applied Mathematics (Fudan University), Shanghai, China

Shuqin Zhang

Roles Conceptualization, Funding acquisition, Project administration, Supervision, Writing – review & editing

zhangs@fudan.edu.cn

Affiliations School of Mathematical Sciences, Fudan University, Shanghai, China, Key Laboratory of Mathematics for Nonlinear Science (Fudan University), Ministry of Education, Shanghai, China, Shanghai Key Laboratory for Contemporary Applied Mathematics (Fudan University), Shanghai, China

Competing Interests

The authors have declared that no competing interests exist.