About the Authors

Takeru Kameda

Roles Conceptualization, Formal analysis, Investigation, Methodology, Software, Visualization, Writing – original draft, Writing – review & editing

Affiliations Graduate School of Science, Hiroshima University, Higashi-Hiroshima, Hiroshima, Japan, RIKEN Center for Biosystems Dynamics Research (BDR), Wako, Saitama, Japan

Katsura Asano

Roles Conceptualization, Formal analysis, Funding acquisition, Project administration, Supervision, Writing – original draft, Writing – review & editing

kasano@ksu.edu (KA); yuichi.togashi@riken.jp (YT)

Affiliations Molecular Cellular and Developmental Biology Program, Division of Biology, Kansas State University, Manhattan, Kansas, United States of America, Graduate School of Integrated Sciences for Life, Hiroshima University, Higashi-Hiroshima, Hiroshima, Japan, Hiroshima Research Center for Healthy Aging (HiHA), Hiroshima University, Higashi-Hiroshima, Hiroshima, Japan

Yuichi Togashi

Roles Conceptualization, Formal analysis, Funding acquisition, Methodology, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

kasano@ksu.edu (KA); yuichi.togashi@riken.jp (YT)

address: College of Life Sciences, Ritsumeikan University, Kusatsu, Shiga, Japan

Affiliations Graduate School of Integrated Sciences for Life, Hiroshima University, Higashi-Hiroshima, Hiroshima, Japan, Research Center for the Mathematics on Chromatin Live Dynamics (RcMcD), Hiroshima University, Higashi-Hiroshima, Hiroshima, Japan, RIKEN Center for Biosystems Dynamics Research (BDR), Higashi-Hiroshima, Hiroshima, Japan

Competing Interests

The authors have declared that no competing interests exist.