About the Authors

Zhiyuan Li

Roles Conceptualization, Data curation, Formal analysis, Methodology, Visualization, Writing – original draft, Writing – review & editing

Affiliations Center for Quantitative Biology, Peking University, Beijing, China, Peking-Tsinghua Center for Life Sciences, Peking University, Beijing, China, Center for the Physics of Biological Function, Princeton University, Princeton, New Jersey, United States of America, Princeton Center for Theoretical Science, Princeton University, Princeton, New Jersey, United States of America

Bo Liu

Roles Data curation, Formal analysis

Affiliation Yuanpei College, Peking University, Beijing, China

Sophia Hsin-Jung Li

Roles Data curation

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Christopher G. King

Roles Data curation

Affiliation Department of Physics, Princeton University, Princeton, New Jersey, United States of America

Zemer Gitai

Roles Conceptualization, Data curation

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Ned S. Wingreen

Roles Conceptualization, Funding acquisition, Supervision, Writing – review & editing

wingreen@princeton.edu

Affiliations Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America, Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.