About the Authors

Yuji Sakai

Roles Conceptualization, Data curation, Investigation, Validation, Visualization, Writing – original draft

Affiliations iTHES Research Group, RIKEN, Wako, Japan, Theoretical Biology Laboratory, RIKEN, Wako, Japan, Department of Biochemistry and Molecular Biology, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Atsushi Mochizuki

Roles Conceptualization, Funding acquisition, Validation, Writing – original draft

Affiliations iTHES Research Group, RIKEN, Wako, Japan, Theoretical Biology Laboratory, RIKEN, Wako, Japan, iTHEMS Program, RIKEN, Wako, Japan, CREST, JST 4-1-8 Honcho, Kawaguchi, Japan

Kazuhisa Kinoshita

Roles Conceptualization, Funding acquisition, Validation, Writing – original draft

Affiliation Chromosome Dynamics Laboratory, RIKEN, Wako, Japan

Tatsuya Hirano

Roles Conceptualization, Supervision, Validation, Writing – original draft

Affiliation Chromosome Dynamics Laboratory, RIKEN, Wako, Japan

Masashi Tachikawa

Roles Conceptualization, Funding acquisition, Investigation, Supervision, Validation, Writing – original draft

mtach@riken.jp

Affiliations iTHES Research Group, RIKEN, Wako, Japan, Theoretical Biology Laboratory, RIKEN, Wako, Japan, iTHEMS Program, RIKEN, Wako, Japan

Competing Interests

The authors have declared that no competing interests exist.