About the Authors

Yan Zhang

Contributed equally to this work with: Yan Zhang, Shantao Li

Affiliations Program in Computational Biology and Bioinformatics, Yale University, New Haven, Connecticut, United States of America, Department of Molecular Biophysics and Biochemistry, Yale University, New Haven, Connecticut, United States of America, Department of Biomedical Informatics, College of Medicine, The Ohio State University, Columbus, Ohio, United States of America

Shantao Li

Contributed equally to this work with: Yan Zhang, Shantao Li

Affiliation Program in Computational Biology and Bioinformatics, Yale University, New Haven, Connecticut, United States of America

Alexej Abyzov

abyzov.alexej@mayo.edu (AA); mark@gersteinlab.org (MBG)

Affiliation Department of Health Sciences Research, Center for Individualized Medicine, Mayo Clinic, Rochester, Minnesota, United States of America

Mark B. Gerstein

abyzov.alexej@mayo.edu (AA); mark@gersteinlab.org (MBG)

Affiliations Program in Computational Biology and Bioinformatics, Yale University, New Haven, Connecticut, United States of America, Department of Molecular Biophysics and Biochemistry, Yale University, New Haven, Connecticut, United States of America, Department of Computer Science, Yale University, New Haven, Connecticut, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: AA YZ SL MBG. Formal analysis: YZ SL. Investigation: YZ SL AA MBG. Methodology: YZ SL AA MBG. Software: YZ SL AA. Supervision: AA MBG. Writing – original draft: YZ SL AA MBG. Writing – review & editing: YZ SL AA MBG.