About the Authors

William R. Holmes

william.holmes@vanderbilt.edu

Affiliation Department of Physics and Astronomy, Vanderbilt University, Nashville, Tennessee, United States of America

JinSeok Park

Affiliation Department of Biomedical Engineering, Yale University, New Haven, Connecticut, United States of America

Andre Levchenko

Affiliation Department of Biomedical Engineering, Yale University, New Haven, Connecticut, United States of America

Leah Edelstein-Keshet

Affiliation Department of Mathematics, University of British Columbia, Vancouver BC, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: WRH AL LEK. Formal analysis: WRH. Funding acquisition: WRH AL LEK. Methodology: WRH LEK. Project administration: WRH LEK. Software: WRH. Supervision: AL LEK. Writing – original draft: WRH JP AL LEK.