About the Authors

Tom Sumner

Current address: London School of Hygiene and Tropical Medicine, London, United Kingdom

Affiliation The Pirbright Institute, Pirbright, Surrey, United Kingdom

Richard J. Orton

Affiliation Boyd Orr Centre for Population and Ecosystem Health, Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Darren M. Green

Affiliation Institute of Aquaculture, University of Stirling, Stirling, Stirlingshire, United Kingdom

Rowland R. Kao

Affiliation Boyd Orr Centre for Population and Ecosystem Health, Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Simon Gubbins

simon.gubbins@pirbright.ac.uk

Affiliation The Pirbright Institute, Pirbright, Surrey, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: SG DMG RRK. Formal analysis: SG TS. Methodology: TS RJO DMG RRK SG. Software: TS SG. Writing – original draft: SG TS. Writing – review & editing: TS RJO DMG RRK SG.