About the Authors

Sophie K. Kay

Affiliation Department of Computer Science, University of Oxford, Oxford, U.K.

Heather A. Harrington

Affiliation Mathematical Institute, University of Oxford, Oxford, U.K.

Sarah Shepherd

† Deceased.

Affiliation School of Mathematical Sciences, University of Nottingham, Nottingham, U.K.

Keith Brennan

Affiliation Wellcome Trust Centre for Cell Matrix Research, University of Manchester, Manchester, U.K.

Trevor Dale

Affiliation School of Biosciences, Cardiff University, Cardiff, U.K.

James M. Osborne

Affiliation School of Mathematics and Statistics, University of Melbourne, Melbourne, Australia

David J. Gavaghan

Affiliation Department of Computer Science, University of Oxford, Oxford, U.K.

Helen M. Byrne

helen.byrne@maths.ox.ac.uk

Affiliations Department of Computer Science, University of Oxford, Oxford, U.K., Mathematical Institute, University of Oxford, Oxford, U.K.

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Formal analysis: SKK HAH KB TD JMO DJG HMB. Investigation: SKK. Methodology: SKK HAH JMO DJG HMB. Resources: SKK HAH SS KB TD JMO DJG HMB. Supervision: JMO DJG HMB. Writing – original draft: SKK HAH KB TD JMO DJG HMB.