About the Authors

Rahul Subramanian

rsfive@uchicago.edu

Affiliation Department of Ecology and Evolution, University of Chicago, Chicago, Illinois, United States of America

Andrea L. Graham

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Bryan T. Grenfell

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Nimalan Arinaminpathy

Affiliation Department of Infectious Disease Epidemiology, Imperial College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NA ALG BTG. Performed the experiments: RS. Analyzed the data: RS ALG NA BTG. Wrote the paper: RS NA ALG BTG.