About the Authors

George L. Sutphin

sutphin@gmail.com (GLS); ron.korstanje@jax.org (RK)

‡ These authors share first authorship on this work.

Affiliation The Jackson Laboratory, Bar Harbor, ME, United States of America

J. Matthew Mahoney

‡ These authors share first authorship on this work.

Affiliation Department of Neurological Sciences, University of Vermont College of Medicine, Burlington, VT, United States of America

Keith Sheppard

Affiliation The Jackson Laboratory, Bar Harbor, ME, United States of America

David O. Walton

Affiliation The Jackson Laboratory, Bar Harbor, ME, United States of America

Ron Korstanje

sutphin@gmail.com (GLS); ron.korstanje@jax.org (RK)

Affiliation The Jackson Laboratory, Bar Harbor, ME, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: GLS JMM RK. Data curation: GLS RK. Formal analysis: GLS JMM. Funding acquisition: RK. Investigation: GLS JMM. Methodology: GLS JMM. Project administration: GLS. Resources: DOW RK. Software: GLS JMM KS DOW. Supervision: RK. Validation: GLS JMM RK. Visualization: GLS JMM. Writing – original draft: GLS JMM. Writing – review & editing: GLS JMM KS DOW RK.