About the Authors

Ryan C. Williamson

Affiliations Center for the Neural Basis of Cognition, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, School of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Machine Learning, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Benjamin R. Cowley

Affiliations Center for the Neural Basis of Cognition, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Machine Learning, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Ashok Litwin-Kumar

Affiliation Center for Theoretical Neuroscience, Columbia University, New York City, New York, United States of America

Brent Doiron

Affiliations Center for the Neural Basis of Cognition, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Mathematics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Adam Kohn

Affiliations Dominick Purpura Department of Neuroscience, Albert Einstein College of Medicine, Bronx, New York, United States of America, Department of Ophthalmology and Vision Sciences, Albert Einstein College of Medicine, Bronx, New York, United States of America, Department of Systems and Computational Biology, Albert Einstein College of Medicine, Bronx, New York, United States of America

Matthew A. Smith

Contributed equally to this work with: Matthew A. Smith, Byron M. Yu

Affiliations Center for the Neural Basis of Cognition, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Ophthalmology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Bioengineering, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Fox Center for Vision Restoration, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Byron M. Yu

Contributed equally to this work with: Matthew A. Smith, Byron M. Yu

byronyu@cmu.edu

Affiliations Center for the Neural Basis of Cognition, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Electrical and Computer Engineering, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Biomedical Engineering, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: RCW BRC ALK BD MAS BMY. Data curation: RCW BRC AK MAS BMY. Formal analysis: RCW BRC MAS BMY. Funding acquisition: RCW BRC ALK BD AK MAS BMY. Investigation: RCW AK MAS BMY. Methodology: RCW BRC ALK BD AK MAS BMY. Project administration: RCW MAS BMY. Resources: RCW ALK BD AK MAS BMY. Software: RCW ALK BMY. Supervision: MAS BMY. Validation: RCW MAS BMY. Visualization: RCW MAS BMY. Writing – original draft: RCW MAS BMY. Writing – review & editing: RCW BRC ALK BD AK MAS BMY.