About the Authors

Bonnie Nijhof

Contributed equally to this work with: Bonnie Nijhof, Anna Castells-Nobau

Affiliation Department of Human Genetics, Donders Institute for Brain, Cognition and Behaviour, Radboud University Medical Center, Nijmegen, the Netherlands

Anna Castells-Nobau

Contributed equally to this work with: Bonnie Nijhof, Anna Castells-Nobau

Affiliation Department of Human Genetics, Donders Institute for Brain, Cognition and Behaviour, Radboud University Medical Center, Nijmegen, the Netherlands

Louis Wolf

Affiliation Microscopical Imaging Centre (MIC), Radboud University Medical Center, Nijmegen, the Netherlands

Jolanda M. Scheffer-de Gooyert

Affiliation Department of Human Genetics, Donders Institute for Brain, Cognition and Behaviour, Radboud University Medical Center, Nijmegen, the Netherlands

Ignacio Monedero

Affiliations Department of Biology, Universidad Autónoma de Madrid, Madrid, Spain, Department of Clinical and Experimental Medicine, Linköping University, Linköping, Sweden

Laura Torroja

Affiliation Department of Biology, Universidad Autónoma de Madrid, Madrid, Spain

Lluis Coromina

Affiliation Research Group on Statistics, Econometrics and Health (GRECS) and CIBER of Epidemiology and Public Health (CIBERESP), University of Girona, Girona, Spain

Jeroen A. W. M. van der Laak

Affiliations Department of Pathology, Radboud University Medical Center, Nijmegen, the Netherlands, Microscopical Imaging Centre (MIC), Radboud University Medical Center, Nijmegen, the Netherlands

Annette Schenck

Annette.Schenck@radboudumc.nl

Affiliation Department of Human Genetics, Donders Institute for Brain, Cognition and Behaviour, Radboud University Medical Center, Nijmegen, the Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BN ACN JAWMvdL AS. Performed the experiments: BN ACN LW JMSdG IM. Analyzed the data: BN ACN. Contributed reagents/materials/analysis tools: LT LC. Wrote the paper: BN ACN JAWMvdL AS.