About the Authors

Chuan Gao

Affiliation Department of Statistical Science, Duke University, Durham, North Carolina, United States of America

Ian C. McDowell

Affiliation Program in Computational Biology and Bioinformatics, Duke University, Durham, North Carolina, United States of America

Shiwen Zhao

Affiliation Program in Computational Biology and Bioinformatics, Duke University, Durham, North Carolina, United States of America

Christopher D. Brown

Affiliation Department of Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Barbara E. Engelhardt

bee@princeton.edu

Affiliation Department of Computer Science, Center for Statistics and Machine Learning, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BEE CG. Performed the experiments: CG. Analyzed the data: CG BEE. Contributed reagents/materials/analysis tools: ICM SZ. Wrote the paper: BEE CG CDB.