About the Authors

Gregory R. Johnson

Affiliations Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Center for Bioimage Informatics, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Jieyue Li

Affiliations Center for Bioimage Informatics, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Biomedical Engineering, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Aabid Shariff

Affiliations Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Center for Bioimage Informatics, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Gustavo K. Rohde

‡ These authors are joint senior authors on this work.

Affiliations Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Center for Bioimage Informatics, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Biomedical Engineering, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Electrical and Computer Engineering, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Robert F. Murphy

murphy@cmu.edu

‡ These authors are joint senior authors on this work.

Affiliations Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Center for Bioimage Informatics, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Biomedical Engineering, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Departments of Biological Sciences and Machine Learning, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Faculty of Biology and Freiburg Institute for Advanced Studies, Albert Ludwig University of Freiburg, Freiburg, Germany

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GRJ JL AS GKR RFM. Performed the experiments: GRJ JL AS. Analyzed the data: GRJ JL AS GKR RFM. Wrote the paper: GRJ JL AS GKR RFM.