About the Authors

Gabriel Koch Ocker

Affiliations Department of Neuroscience, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Center for the Neural Basis of Cognition, University of Pittsburgh and Carnegie Melon University, Pittsburgh, Pennsylvania, United States of America

Ashok Litwin-Kumar

Affiliations Center for the Neural Basis of Cognition, University of Pittsburgh and Carnegie Melon University, Pittsburgh, Pennsylvania, United States of America, Department of Mathematics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Center for Theoretical Neuroscience, Columbia University, New York, New York, United States of America

Brent Doiron

bdoiron@pitt.edu

Affiliations Center for the Neural Basis of Cognition, University of Pittsburgh and Carnegie Melon University, Pittsburgh, Pennsylvania, United States of America, Department of Mathematics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GKO ALK BD. Performed the experiments: GKO. Analyzed the data: GKO. Wrote the paper: GKO ALK BD.