About the Authors

Shuai Yuan

Affiliation Mathematics & Computer Science Department, Emory University, Atlanta, Georgia, United States of America

H. Richard Johnston

Affiliation Department of Biostatistics and Bioinformatics, Rollins School of Public Health, Emory University, Atlanta, Georgia, United States of America

Guosheng Zhang

Affiliation Department of Genetics, Department of Biostatistics, Department of Computer Science, University of North Carolina, Chapel Hill, Chapel Hill, North Carolina, United States of America

Yun Li

Affiliation Department of Genetics, Department of Biostatistics, Department of Computer Science, University of North Carolina, Chapel Hill, Chapel Hill, North Carolina, United States of America

Yi-Juan Hu

Affiliation Department of Biostatistics and Bioinformatics, Rollins School of Public Health, Emory University, Atlanta, Georgia, United States of America

Zhaohui S. Qin

zhaohui.qin@emory.edu

Affiliation Department of Biostatistics and Bioinformatics, Rollins School of Public Health, Emory University, Atlanta, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZSQ SY. Performed the experiments: SY GZ. Analyzed the data: ZSQ SY YH YL. Contributed reagents/materials/analysis tools: YH YL. Wrote the paper: SY ZSQ HRJ YH YL.