About the Authors

Daniel K. Wells

Affiliations Department of Engineering Sciences and Applied Mathematics, Northwestern University, Evanston, Illinois, United States of America, Northwestern University Physical Sciences-Oncology Center, Evanston, Illinois, United States of America

Yishan Chuang

Affiliation Department of Chemical and Biological Engineering, Northwestern University, Evanston, Illinois, United States of America

Louis M. Knapp

Affiliation Department of Chemical and Biological Engineering, Northwestern University, Evanston, Illinois, United States of America

Dirk Brockmann

Current address: Robert Koch Institute, Berlin, Germany

Affiliations Department of Engineering Sciences and Applied Mathematics, Northwestern University, Evanston, Illinois, United States of America, Northwestern University Physical Sciences-Oncology Center, Evanston, Illinois, United States of America, Robert H. Lurie Comprehensive Cancer Center, Northwestern University, Evanston, Illinois, United States of America, Northwestern Institute on Complex Science, Northwestern University, Evanston, Illinois, United States of America, Institute for Theoretical Biology, Humboldt University Berlin, Berlin, Germany

William L. Kath

Affiliations Department of Engineering Sciences and Applied Mathematics, Northwestern University, Evanston, Illinois, United States of America, Northwestern University Physical Sciences-Oncology Center, Evanston, Illinois, United States of America, Robert H. Lurie Comprehensive Cancer Center, Northwestern University, Evanston, Illinois, United States of America, Northwestern Institute on Complex Science, Northwestern University, Evanston, Illinois, United States of America, Chemistry of Life Processes Institute, Northwestern University, Evanston, Illinois, United States of America

Joshua N. Leonard

j-leonard@northwestern.edu

Affiliations Northwestern University Physical Sciences-Oncology Center, Evanston, Illinois, United States of America, Department of Chemical and Biological Engineering, Northwestern University, Evanston, Illinois, United States of America, Robert H. Lurie Comprehensive Cancer Center, Northwestern University, Evanston, Illinois, United States of America, Chemistry of Life Processes Institute, Northwestern University, Evanston, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DKW YC LMK DB WLK JNL. Performed the experiments: DKW. Analyzed the data: DKW WLK JNL. Wrote the paper: DKW YC JNL.