About the Authors

Oren E. Livne

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Lide Han

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Gorka Alkorta-Aranburu

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

William Wentworth-Sheilds

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Mark Abney

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Carole Ober

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Dan L. Nicolae

nicolae@galton.uchicago.edu

Affiliations Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America, Departments of Medicine, and Statistics, The University of Chicago, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: OEL MA CO DLN. Performed the experiments: OEL LH WWS. Analyzed the data: OEL MA CO DLN. Contributed reagents/materials/analysis tools: OEL LH GAA WWS. Wrote the paper: OEL GAA MA CO DLN.