About the Authors

Luca Giuggioli

Affiliations Bristol Centre for Complexity Sciences, University of Bristol, Bristol, United Kingdom, School of Biological Sciences, University of Bristol, Bristol, United Kingdom, Department of Engineering Mathematics, University of Bristol, Bristol, United Kingdom

Thomas J. McKetterick

Affiliations Bristol Centre for Complexity Sciences, University of Bristol, Bristol, United Kingdom, School of Biological Sciences, University of Bristol, Bristol, United Kingdom

Marc Holderied

marc.holderied@bristol.ac.uk

Affiliation Department of Engineering Mathematics, University of Bristol, Bristol, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LG MH. Performed the experiments: LG MH. Analyzed the data: TJM LG MH. Contributed reagents/materials/analysis tools: MH LG. Wrote the paper: LG MH.