About the Authors

Adrian J. de Froment

adriandefroment@gmail.com

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Daniel I. Rubenstein

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Simon A. Levin

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: AJdF DIR SAL. Conceived of and designed the model: AJdF DIR SAL. Programmed and analysed the model: AJdF.