About the Authors

Penelope A. Hancock

penelope.hancock@zoo.ox.ac.uk

Affiliations Department of Zoology, University of Oxford, Oxford, United Kingdom, School of Life Sciences, University of Warwick, Coventry, United Kingdom

Yasmin Rehman

Affiliation Field Epidemiology Service (Birmingham), Public Health England, West Midlands, United Kingdom

Ian M. Hall

Affiliation Emergency Response Unit, Public Health England, Porton Down, United Kingdom

Obaghe Edeghere

Affiliation Field Epidemiology Service (Birmingham), Public Health England, West Midlands, United Kingdom

Leon Danon

Affiliations School of Mathematics, Queen Mary University of London, London, United Kingdom, Warwick Mathematics Institute, University of Warwick, Coventry, United Kingdom

Thomas A. House

Affiliation Warwick Mathematics Institute, University of Warwick, Coventry, United Kingdom

Matthew J. Keeling

Affiliations School of Life Sciences, University of Warwick, Coventry, United Kingdom, Warwick Mathematics Institute, University of Warwick, Coventry, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PAH LD TAH MJK. Performed the experiments: PAH. Analyzed the data: PAH. Contributed reagents/materials/analysis tools: YR IMH OE. Wrote the paper: PAH MJK.