About the Authors

Marko Jusup

mjusup@gmail.com

Affiliation Department of Biology, Kyushu University, Fukuoka, Japan

Tadasu Matsuo

Affiliation Faculty of Economics, Ritsumeikan University, Kusatsu, Japan

Yoh Iwasa

Affiliation Department of Biology, Kyushu University, Fukuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: MJ YI. Developed the basic model: TM YI. Generalized the model: MJ. Ran numerical simulations: MJ. Discussed the results and implications: MJ TM YI.