About the Authors

Marcus J. Tindall

m.tindall@reading.ac.uk (MJT); a.clerk@reading.ac.uk (AC)

Affiliations Department of Mathematics & Statistics, University of Reading, Reading, Berkshire, United Kingdom, School of Biological Sciences, University of Reading, Reading, Berkshire, United Kingdom

Angela Clerk

m.tindall@reading.ac.uk (MJT); a.clerk@reading.ac.uk (AC)

Affiliation School of Biological Sciences, University of Reading, Reading, Berkshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MJT AC. Wrote the paper: MJT AC. Analyzed the experimental data: AC. Formulated and solved mathematical models: MJT. Informed and calculated mathematical model parameter values: MJT.