About the Authors

Shyam Srinivasan

Contributed equally to this work with: Shyam Srinivasan, Jia Sheng Hu

shyams@uci.edu (SS); adlander@uci.edu (ADL); emonuki@uci.edu (ESM)

Affiliations Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America, Center for Complex Biological Systems, University of California, Irvine, California, United States of America, Department of Computer Science, University of California, Irvine, California, United States of America

Jia Sheng Hu

Contributed equally to this work with: Shyam Srinivasan, Jia Sheng Hu

Affiliation Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America

D. Spencer Currle

Affiliations Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America, Department of Developmental Neurobiology, St Jude Children's Research Hospital, Memphis, Tennessee, United States of America

Ernest S. Fung

Affiliation Department of Pathology and Laboratory Medicine, University of California, Irvine, California, United States of America

Wayne B. Hayes

Affiliation Department of Computer Science, University of California, Irvine, California, United States of America

Arthur D. Lander

shyams@uci.edu (SS); adlander@uci.edu (ADL); emonuki@uci.edu (ESM)

Affiliations Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America, Center for Complex Biological Systems, University of California, Irvine, California, United States of America

Edwin S. Monuki

shyams@uci.edu (SS); adlander@uci.edu (ADL); emonuki@uci.edu (ESM)

Affiliations Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America, Center for Complex Biological Systems, University of California, Irvine, California, United States of America, Department of Pathology and Laboratory Medicine, University of California, Irvine, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS JSH ESF DSC WBH ADL ESM. Performed the experiments: SS JSH ESF DSC. Analyzed the data: SS JSH ADL ESM. Wrote the paper: SS JSH ADL ESM. Software for simulations and analysis of models: SS.