About the Authors

Eric A. Zilli

zilli@bu.edu

Affiliation Department of Psychology, Boston University, Boston, Massachusetts, United States of America

Motoharu Yoshida

Affiliation Department of Psychology, Boston University, Boston, Massachusetts, United States of America

Babak Tahvildari

Current address: Department of Neurobiology, School of Medicine, Yale University, New Haven, Connecticut, United States of America

Affiliation Department of Neurology and Neurosurgery, Montreal Neurological Institute, McGill University, Montreal, Canada

Lisa M. Giocomo

Current address: Kavli Institute for Systems Neuroscience and Centre for the Biology of Memory, Trondheim, Norway

Affiliation Department of Psychology, Boston University, Boston, Massachusetts, United States of America

Michael E. Hasselmo

Affiliation Department of Psychology, Boston University, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MY BT LMG MEH. Performed the experiments: MY BT LMG. Analyzed the data: MY BT LMG. Wrote the paper: EAZ MY BT LMG. Programmed and carried out the simulations and performed the mathematical analyses: EAZ. Programmed the simulations: MEH.