About the Authors

A. Aldo Faisal

To whom correspondence should be addressed. E-mail: a.faisal@zoo.cam.ac.uk

Affiliation Department of Zoology, Cambridge University, Cambridge, United Kingdom

Simon B Laughlin

Affiliation Department of Zoology, Cambridge University, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

AAF and SBL conceived the problem. AAF implemented the software, developed models and protocols, carried out the simulation, analyzed the data, and interpreted the results. AAF wrote the paper with SBL. The Modigliani simulation software was developed by AAF as part of his Ph.D.