About the Authors

Fusako Kitaura

Roles Investigation

Affiliation Chromatin Biochemistry Research Group, Combined Program on Microbiology and Immunology, Research Institute for Microbial Diseases, Osaka University, Yamadaoka, Suita, Osaka, Japan

Miyuki Yuno

Roles Investigation

Affiliation Chromatin Biochemistry Research Group, Combined Program on Microbiology and Immunology, Research Institute for Microbial Diseases, Osaka University, Yamadaoka, Suita, Osaka, Japan

Toshitsugu Fujita

Roles Funding acquisition, Investigation, Writing – review & editing

Affiliations Chromatin Biochemistry Research Group, Combined Program on Microbiology and Immunology, Research Institute for Microbial Diseases, Osaka University, Yamadaoka, Suita, Osaka, Japan, Department of Biochemistry and Genome Biology, Hirosaki University Graduate School of Medicine, Zaifu-cho, Hirosaki, Aomori, Japan

Shigeharu Wakana

Roles Investigation

Affiliations Technology and Development Team for Mouse Phenotype Analysis, RIKEN BioResource Research Center, Koyadai, Tsukuba, Ibaraki, Japan, Department of Gerontology, Institute of Biomedical Research and Innovation, Minatojima Minamiachi, Chuo-ku, Kobe, Japan

Jun Ueda

Roles Investigation

Affiliations Center for Genetic Analysis of Biological Responses, Research Institute for Microbial Diseases, Osaka University, Yamadaoka, Suita, Japan, Center for Advanced Research and Education (CARE), Asahikawa Medical University, Asahikawa, Japan

Kazuo Yamagata

Roles Investigation

Affiliations Center for Genetic Analysis of Biological Responses, Research Institute for Microbial Diseases, Osaka University, Yamadaoka, Suita, Japan, Faculty of Biology-Oriented Science and Technology, Kindai University, Nishimitani, Kinokawa City, Wakayama, Japan

Hodaka Fujii

Roles Conceptualization, Funding acquisition, Investigation, Supervision, Writing – original draft, Writing – review & editing

hodaka@hirosaki-u.ac.jp

Affiliations Chromatin Biochemistry Research Group, Combined Program on Microbiology and Immunology, Research Institute for Microbial Diseases, Osaka University, Yamadaoka, Suita, Osaka, Japan, Department of Biochemistry and Genome Biology, Hirosaki University Graduate School of Medicine, Zaifu-cho, Hirosaki, Aomori, Japan

Competing Interests

I have read the journal's policy and the authors of this manuscript have the following competing interests: T.F. and H.F. are founders and directors of Epigeneron, Inc. H.F. is an Academic Editor of PLOS ONE. This does not alter our adherence to PLOS ONE policies on sharing data and materials.