About the Authors

Noboru Sasaki

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Writing – original draft

Affiliation Department of Clinical Sciences, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Kazuhiro Ishi

Roles Investigation

Affiliation Department of Clinical Sciences, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Nobuki Kudo

Roles Methodology, Resources

Affiliation Division of Bioengineering and Bioinformatics, Graduate School of Information Science and Technology, Hokkaido University, Sapporo, Japan

Shouta M. M. Nakayama

Roles Investigation, Validation

Affiliation Department of Environmental Veterinary Sciences, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Kensuke Nakamura

Roles Writing – review & editing

Current address: Organization for Promotion of Tenure Track, University of Miyazaki, Miyazaki, Japan

Affiliation Veterinary Teaching Hospital, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Keitaro Morishita

Roles Writing – review & editing

Affiliation Veterinary Teaching Hospital, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Hiroshi Ohta

Roles Supervision, Writing – review & editing

Affiliation Department of Clinical Sciences, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Mayumi Ishizuka

Roles Investigation

Affiliation Department of Environmental Veterinary Sciences, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Mitsuyoshi Takiguchi

Roles Conceptualization, Funding acquisition, Supervision

mtaki@vetmed.hokudai.ac.jp

Affiliation Department of Clinical Sciences, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.