About the Authors

Takashi Ikeda

wild_wolf_traveler@yahoo.co.jp

Affiliation Faculty of Environmental Earth Science, Hokkaido University, Sapporo, Hokkaido, Japan

Kenta Uchida

Affiliation Graduate School of Environmental Earth Science, Hokkaido University, Sapporo, Hokkaido, Japan

Yukiko Matsuura

Affiliation Hokkaido Research Center, Forestry and Forest Products Research Institute, Sapporo, Hokkaido, Japan

Hiroshi Takahashi

Affiliation Kansai Research Center, Forestry and Forest Products Research Institute, Kyoto, Kyoto, Japan

Tsuyoshi Yoshida

Affiliation Wildlife Management Laboratory, Rakuno Gakuen University, Ebetsu, Hokkaido, Japan

Koichi Kaji

Affiliation Laboratory of Wildlife Management, Tokyo University of Agriculture and Technology, Fuchu, Tokyo, Japan

Itsuro Koizumi

Affiliations Faculty of Environmental Earth Science, Hokkaido University, Sapporo, Hokkaido, Japan, Graduate School of Environmental Earth Science, Hokkaido University, Sapporo, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: TI KU IK. Data curation: TI. Formal analysis: TI IK. Funding acquisition: TI TY. Investigation: TI YM HT. Methodology: TI KU IK. Project administration: TI TY KK IK. Resources: TI. Software: TI. Supervision: TI KU IK. Validation: TI KU YM HT TY KK IK. Visualization: TI. Writing – original draft: TI KU YM HT TY KK IK. Writing – review & editing: TI IK.