About the Authors

Ahmed Siah

ahmed.siah@cahs-bc.ca

Affiliation British Columbia Centre for Aquatic Health Sciences, Campbell River, British Columbia, Canada

Diane B. Morrison

Affiliation Marine Harvest Canada, Campbell River, British Columbia, Canada

Elena Fringuelli

Affiliation Veterinary Sciences Division, AFBI Stormont, Stoney Road, Belfast, United Kingdom

Paul Savage

Affiliation Veterinary Sciences Division, AFBI Stormont, Stoney Road, Belfast, United Kingdom

Zina Richmond

Affiliation British Columbia Centre for Aquatic Health Sciences, Campbell River, British Columbia, Canada

Robert Johns

Affiliation British Columbia Centre for Aquatic Health Sciences, Campbell River, British Columbia, Canada

Maureen K. Purcell

Affiliation US Geological Survey, Western Fisheries Research Center, Seattle, WA, United States of America

Stewart C. Johnson

Affiliation Pacific Biological Station, Nanaimo, British Columbia, Canada

Sonja M. Saksida

Affiliation British Columbia Centre for Aquatic Health Sciences, Campbell River, British Columbia, Canada

Competing Interests

This study was funded by Marine Harvest Canada. This commercial affiliation does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AS DBM SMS. Performed the experiments: DBM SMS SCJ MKP. Analyzed the data: AS EF PS MKP SCJ. Contributed reagents/materials/analysis tools: DBM SMS EF RJ ZR MKP SCJ. Wrote the paper: AS DBM EF PS SMS MKP.