About the Authors

Clarisse Lorin

clarisse.m.lorin@gsk.com

Affiliation GSK Vaccines, Rixensart, Belgium

Yannick Vanloubbeeck

Affiliation GSK Vaccines, Rixensart, Belgium

Sébastien Baudart

Affiliation GSK Vaccines, Rixensart, Belgium

Michaël Ska

Affiliation GSK Vaccines, Rixensart, Belgium

Babak Bayat

Affiliation GSK Vaccines, Rixensart, Belgium

Geoffroy Brauers

Affiliation GSK Vaccines, Rixensart, Belgium

Géraldine Clarinval

Affiliation GSK Vaccines, Rixensart, Belgium

Marie-Noëlle Donner

Affiliation GSK Vaccines, Rixensart, Belgium

Martine Marchand

Affiliation GSK Vaccines, Rixensart, Belgium

Marguerite Koutsoukos

Affiliation GSK Vaccines, Rixensart, Belgium

Pascal Mettens

Affiliation GSK Vaccines, Rixensart, Belgium

Joe Cohen

Affiliation GSK Vaccines, Rixensart, Belgium

Gerald Voss

Affiliation GSK Vaccines, Rixensart, Belgium

Competing Interests

All authors declared that the following interests are relevant to the submitted work: All authors are, or were at the time of the study, employees of the GSK group of companies. JC now works as an independent consultant for GSK Vaccines. CL, YV, MK, GV, JC and PM report ownership of GSK shares. JC, CL, MK and GV are listed as inventors on patents owned by GSK. The F4 construct is disclosed in patent WO2006/013106: “Vaccine for prevention and treatment of HIV-infection”. This does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CL YV MK GV JC BB PM. Performed the experiments: SB MS GB GC BB MND. Analyzed the data: CL YV BB. Wrote the paper: CL YV SB MS BB GB GC MND MM MK PM JC GV.