About the Authors

Ileana Soto

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Gareth R. Howell

Affiliations The Jackson Laboratory, Bar Harbor, Maine, United States of America, The Howard Hughes Medical Institute, Bar Harbor, Maine, United States of America

Cai W. John

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Joseph L. Kief

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Richard T. Libby

Affiliation Departments of Ophthalmology and Biomedical Genetics, University of Rochester Medical Center, Rochester, New York, United States of America

Simon W. M. John

simon.john@jax.org

Affiliations The Jackson Laboratory, Bar Harbor, Maine, United States of America, The Howard Hughes Medical Institute, Bar Harbor, Maine, United States of America, Department of Ophthalmology, Tufts University of Medicine, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SWMJ IS GRH. Performed the experiments: IS CWJ JLK. Analyzed the data: IS GRH RTL. Contributed to the writing of the manuscript: IS GRH RTL SWMJ.