About the Authors

Yan Zhou

Affiliation School of Biological Sciences, Nanyang Technological University, Singapore, Singapore

Wanjin Hong

Affiliation Institute of Molecular and Cell Biology, Singapore, Singapore

Lei Lu

lulei@ntu.edu.sg

Affiliation School of Biological Sciences, Nanyang Technological University, Singapore, Singapore

Competing Interests

Co-author Wanjin Hong is a PLOS ONE Editorial Board member and this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LL. Performed the experiments: YZ LL. Analyzed the data: YZ LL. Contributed reagents/materials/analysis tools: YZ LL. Wrote the paper: LL WH.