About the Authors

Chengmin Wang

Contributed equally to this work with: Chengmin Wang, Bin Wu, Shelan Liu

Affiliation National Research Center For Wildlife Born Diseases, Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Yanyu Zhang

Affiliation Beijing Institute of Transfusion Medicine, Academy of Military Medicine Sciences, Beijing, P.R China

Bin Wu

Contributed equally to this work with: Chengmin Wang, Bin Wu, Shelan Liu

Affiliation National Research Center For Wildlife Born Diseases, Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Shelan Liu

Contributed equally to this work with: Chengmin Wang, Bin Wu, Shelan Liu

Affiliation Department of Infectious Diseases, Zhejiang Center for Disease Control and Prevention, Binjiang District, Hangzhou, Zhejiang Province, China

Ping Xu

Affiliation National Research Center For Wildlife Born Diseases, Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Yanmin Lu

Affiliation National Research Center For Wildlife Born Diseases, Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Jing Luo

Affiliation National Research Center For Wildlife Born Diseases, Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Dale Louis Nolte

Affiliation National Wildlife Research Center, USDA/APHIS/Wildlife Services, United States Department of Agriculture, Fort Collins, Colorado, United States of America

Thomas Jude Deliberto

Affiliation National Wildlife Research Center, USDA/APHIS/Wildlife Services, United States Department of Agriculture, Fort Collins, Colorado, United States of America

Mingxing Duan

Affiliation State Key Laboratory of Biomembrane & Membrane Biotechnology, School of Life Sciences, Tsinghua University, Beijing, China

Hong Zhang

Affiliation National Research Center For Wildlife Born Diseases, Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Hongxuan He

hehx@ioz.ac.cn

Affiliation National Research Center For Wildlife Born Diseases, Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Collected virus sequences: SLL YML PX. Analyzed selection pressure and made 3D picture: CMW BW. Revised the paper: JL DLN HZ MXD TJD. Conceived and designed the experiments: HXH CMW BW HZ. Performed the experiments: CMW BW SLL YML PX. Analyzed the data: BW YYZ CMW. Wrote the paper: CMW HZ HXH.