About the Authors

Chiaki Maruta

Affiliation Laboratory of Neuroscience, Faculty of Pharmaceutical Sciences, Hokkaido University, Sapporo, Japan

Yuhki Saito

Affiliation Laboratory of Neuroscience, Faculty of Pharmaceutical Sciences, Hokkaido University, Sapporo, Japan

Saori Hata

Affiliation Laboratory of Neuroscience, Faculty of Pharmaceutical Sciences, Hokkaido University, Sapporo, Japan

Naoya Gotoh

Affiliation Laboratory of Neuroscience, Faculty of Pharmaceutical Sciences, Hokkaido University, Sapporo, Japan

Toshiharu Suzuki

tsuzuki@pharm.hokudai.ac.jp (TS); yamamoto@pharm.hokudai.ac.jp (TY)

Affiliation Laboratory of Neuroscience, Faculty of Pharmaceutical Sciences, Hokkaido University, Sapporo, Japan

Tohru Yamamoto

tsuzuki@pharm.hokudai.ac.jp (TS); yamamoto@pharm.hokudai.ac.jp (TY)

Affiliation Laboratory of Neuroscience, Faculty of Pharmaceutical Sciences, Hokkaido University, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CM YS TS TY. Performed the experiments: CM YS SH NG. Analyzed the data: CM YS SH NG TS TY. Contributed reagents/materials/analysis tools: CM YS SH TS TY. Wrote the paper: TS TY.