About the Authors

Robert S. Illingworth

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine at the University of Edinburgh, Edinburgh, United Kingdom

Catherine H. Botting

Affiliation BMS Mass Spectrometry and Proteomics Facility, Biomedical Sciences Research Complex, University of St. Andrews, St. Andrews, Fife, United Kingdom

Graeme R. Grimes

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine at the University of Edinburgh, Edinburgh, United Kingdom

Wendy A. Bickmore

W.Bickmore@hgu.mrc.ac.uk (WB); Ragnhild.Eskeland@imbv.uio.no (RE)

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine at the University of Edinburgh, Edinburgh, United Kingdom

Ragnhild Eskeland

W.Bickmore@hgu.mrc.ac.uk (WB); Ragnhild.Eskeland@imbv.uio.no (RE)

Current address: Department of Molecular Biosciences, University of Oslo, Oslo, Norway

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine at the University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RI WB RE. Performed the experiments: RI CB RE. Analyzed the data: RI GG RE. Wrote the paper: RI WB RE.