About the Authors

Alison J. Peel

alisonpeel@gmail.com

Affiliations Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom, Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Kate S. Baker

Affiliations Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom, Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Gary Crameri

Affiliation CSIRO Livestock Industries, Australian Animal Health Laboratory, Geelong, Victoria, Australia

Jennifer A. Barr

Affiliation CSIRO Livestock Industries, Australian Animal Health Laboratory, Geelong, Victoria, Australia

David T. S. Hayman

Affiliations Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom, Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom, Animal Health and Veterinary Laboratories Agency, Wildlife Zoonoses and Vector-Borne Diseases Research Group, Department of Virology, Veterinary Laboratories Agency, Weybridge, New Haw, Addlestone, Surrey, United Kingdom, Department of Biology, Colorado State University, Fort Collins, Colorado, United States of America

Edward Wright

Affiliations School of Life Sciences, University of Westminster, London, United Kingdom, Division of Infection and Immunity, University College London, London, United Kingdom

Christopher C. Broder

Affiliation Department of Microbiology and Immunology, Uniformed Services University, Bethesda, Maryland, United States of America

Andrés Fernández-Loras

Affiliation Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Anthony R. Fooks

Affiliations Animal Health and Veterinary Laboratories Agency, Wildlife Zoonoses and Vector-Borne Diseases Research Group, Department of Virology, Veterinary Laboratories Agency, Weybridge, New Haw, Addlestone, Surrey, United Kingdom, National Centre for Zoonosis Research, Leahurst, Neston, South Wirral, United Kingdom

Lin-Fa Wang

Affiliation CSIRO Livestock Industries, Australian Animal Health Laboratory, Geelong, Victoria, Australia

Andrew A. Cunningham

Affiliation Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

James L. N. Wood

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Competing Interests

Anthony R. Fooks is an Academic Editor for PLoS ONE. Lin-Fa Wang, Gary Crameri and Jennifer Barr are employed by CSIRO Livestock Industries and contributed time and materials to this research. This does not alter the authors′ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AJP AAC JLNW. Performed the experiments: AJP KSB GC DTSH JAB EW. Analyzed the data: AJP KSB GC DTSH JAB EW. Contributed reagents/materials/analysis tools: CCB EW ARF L-FW AAC JLNW. Wrote the paper: AJP KSB GC JAB DTSH EW CCB ARF L-FW AAC JLNW. Collected samples: AJP AFL.