About the Authors

Yuki Kondo

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Software, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

youkikondou@gmail.com

Affiliations Laboratory of Animal Sociology, Department of Biology, Graduate School of Science, Osaka Metropolitan University, Osaka, Japan, Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Osaka, Japan

Kotori Okamoto

Roles Data curation, Formal analysis, Investigation, Methodology

Affiliation Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Osaka, Japan

Yuto Kitamukai

Roles Data curation, Formal analysis

Affiliation Laboratory of Animal Sociology, Department of Biology, Graduate School of Science, Osaka Metropolitan University, Osaka, Japan

Yasunori Koya

Roles Conceptualization, Project administration, Writing – review & editing

Affiliation Department of Biology, Faculty of Education, Gifu University, Gifu, Japan

Satoshi Awata

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Supervision, Writing – review & editing

Affiliations Laboratory of Animal Sociology, Department of Biology, Graduate School of Science, Osaka Metropolitan University, Osaka, Japan, Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.