About the Authors

Sajid Javed

Contributed equally to this work with: Sajid Javed, Leanne Marsay, Karen E. Kempsell

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Leanne Marsay

Contributed equally to this work with: Sajid Javed, Leanne Marsay, Karen E. Kempsell

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Alice Wareham

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Kuiama S. Lewandowski

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Ann Williams

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Michael J. Dennis

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Sally Sharpe

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Richard Vipond

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Nigel Silman

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Graham Ball

Affiliation School of Science and Technology, Nottingham Trent University, Clifton Lane, Nottingham, United Kingdom

Karen E. Kempsell

Contributed equally to this work with: Sajid Javed, Leanne Marsay, Karen E. Kempsell

karen.kempsell@phe.gov.uk

Affiliation Public Health England, Infection Services, Health Protection Agency Porton, Porton Down, Salisbury, Wiltshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KEK SJ LM KSL. Performed the experiments: SJ LM A. Wareham KSL. Analyzed the data: SJ LM A. Wareham KL GB KEK. Contributed reagents/materials/analysis tools: SS MJD A. Williams GB KEK. Wrote the paper: SJ LM GB KEK. Funding: NS. Contributed Resources: RV.