About the Authors

Cheryl A. Mills

Affiliations Centre for Ecology and Conservation, College of Life and Environmental Sciences, University of Exeter, Cornwall Campus, Penryn, Cornwall, United Kingdom, School of Life Sciences, Keele University, Keele, Staffordshire, United Kingdom

Brendan J. Godley

Affiliation Centre for Ecology and Conservation, College of Life and Environmental Sciences, University of Exeter, Cornwall Campus, Penryn, Cornwall, United Kingdom

David J. Hodgson

D.J.Hodgson@exeter.ac.uk

Affiliation Centre for Ecology and Conservation, College of Life and Environmental Sciences, University of Exeter, Cornwall Campus, Penryn, Cornwall, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CAM DJH BJG. Performed the experiments: CAM. Analyzed the data: CAM DJH BJG. Contributed reagents/materials/analysis tools: CAM DJH BJG. Wrote the paper: CAM DJH BJG.