About the Authors

Hiroshi Katoh

Contributed equally to this work with: Hiroshi Katoh, Hiromitsu Inoue

katohh@cc.utsunomiya-u.ac.jp

Current address: Center for Bioscience Research and Education, Utsunomiya University, Mine-machi 350, Utsunomiya, Tochigi 321–8505, Japan

Affiliation NARO Institute of Fruit Tree Science, Fujimoto 2-1, Tsukuba, Ibaraki 305-8605, Japan

Hiromitsu Inoue

Contributed equally to this work with: Hiroshi Katoh, Hiromitsu Inoue

Affiliation Kuchinotsu Citrus Research Station, NARO Institute of Fruit Tree Science, Minami-shimabara, Nagasaki 859–2501, Japan

Toru Iwanami

Current address: Apple Research Center, NARO Institute of Fruit Tree Science, Morioka, Iwate 020-0123, Japan

Affiliation NARO Institute of Fruit Tree Science, Fujimoto 2-1, Tsukuba, Ibaraki 305-8605, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HI TI. Performed the experiments: HK. Analyzed the data: HK. Contributed reagents/materials/analysis tools: HK HI. Wrote the paper: HK.