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Contributed equally to this work with:
* E-mail: email@example.com
Current address: Center for Bioscience Research and Education, Utsunomiya University, Mine-machi 350, Utsunomiya, Tochigi 321–8505, Japan
NARO Institute of Fruit Tree Science, Fujimoto 2-1, Tsukuba, Ibaraki 305-8605, Japan
Kuchinotsu Citrus Research Station, NARO Institute of Fruit Tree Science, Minami-shimabara, Nagasaki 859–2501, Japan
Current address: Apple Research Center, NARO Institute of Fruit Tree Science, Morioka, Iwate 020-0123, Japan
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