About the Authors

Xu Shen

Affiliations Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou, Guangdong, China, Guangdong Provincial Key Lab of Agro-Animal Genomics and Molecular Breeding, Guangzhou, China

Hua Zeng

Affiliation Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou, Guangdong, China

Liang Xie

Affiliations Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou, Guangdong, China, Institute of Animal Science and Veterinary, Hainan Academy of Agricultural Sciences, Haikou, Hainan, China

Jun He

Affiliations Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou, Guangdong, China, Guangdong Provincial Key Lab of Agro-Animal Genomics and Molecular Breeding, Guangzhou, China

Jian Li

Affiliations Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou, Guangdong, China, Guangdong Provincial Key Lab of Agro-Animal Genomics and Molecular Breeding, Guangzhou, China

Xiujuan Xie

Affiliations Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou, Guangdong, China, Guangdong Provincial Key Lab of Agro-Animal Genomics and Molecular Breeding, Guangzhou, China

Chenglong Luo

Affiliation Institute of Animal Science, Guangdong Academy of Agricultural Sciences, Guangzhou, Guangdong, China

Haiping Xu

Affiliations Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou, Guangdong, China, Guangdong Provincial Key Lab of Agro-Animal Genomics and Molecular Breeding, Guangzhou, China

Min Zhou

Affiliation Biotechnology Institute, Jiang Xi Education College, Nanchang, Jiangxi, China

Qinghua Nie

Affiliations Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou, Guangdong, China, Guangdong Provincial Key Lab of Agro-Animal Genomics and Molecular Breeding, Guangzhou, China

Xiquan Zhang

xqzhang@scau.edu.cn

Affiliations Department of Animal Genetics, Breeding and Reproduction, College of Animal Science, South China Agricultural University, Guangzhou, Guangdong, China, Guangdong Provincial Key Lab of Agro-Animal Genomics and Molecular Breeding, Guangzhou, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XS HZ QN XZ. Performed the experiments: XS JH JL XX. Analyzed the data: CL LX HX MZ. Wrote the paper: XS XZ.