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Table 1.

Estimates for the negative binomial parasite aggregation parameter, .

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Figure 1.

The relationship between mean intensity and prevalence.

A The relationship between the mean intensity of infection, , the prevalence of infection, , and the negative binomial aggregation parameter, as described by the relationship in equation 1. B Relationship between the prevalence and intensity of infection as observed in a study of A. lumbricoides [28]. The solid line is the predicted relationship between mean prevalence of infection and worm burden described in equation 1 and plotted in A fitted to estimate the aggregation parameter, = 0.194.

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Table 2.

Percentage of population aged 5 to 14 years in 2011 [40].

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Figure 2.

School attendance for a selection of countries.

This figure was generated by data published by UNICEF for 2005–2010 [41].. For each country there is net attendance rate at primary, in urban (open circles) and rural areas (closed circles) and a net attendance rate for secondary schools (filled squares).

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Table 6.

Percentage of female children who are enrolled in school [45].

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Table 3.

Published estimates of the basic reproductive number for various helminths.

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Table 4.

Published estimates of parasite life expectancy, , in the human host [18], [27].

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Figure 3.

The proportion of A. lumbricoides worms in children aged 5–14, calculated from equation 3.

The demography of the population, A, results in a proportion of 18.4% of the population aged 5–14 years old [45]. Combining this distribution with B the distribution of worms per person by age [57] gives 49.5% of worms in the school-aged group.

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Figure 4.

The proportion of hookworm eggs deposited by children aged 5–14, calculated from equation 5.

The demography of the population, A, gives a proportion of 31.2% of the population aged 5–14 years old [45]. Combining this distribution with B the distribution of egg output by age [36] gives 15.7% of worms in the school-aged group.

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Table 5.

Fraction of worm population or egg output in 5–14 year olds (equations 3 and 5).

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Figure 5.

Critical fraction of the population to be treated.

The predicted relationship between the critical fraction of the human population to be treated, , per annum with efficacy, , 0.9, and the basic reproductive number, , and parasite life expectancy, in years (from equation 11 in the main text).

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Figure 6.

Impact of fraction treated on worm burden, prevalence and effective reproduction number.

The impact of the fraction of the population treated, , on A the mean worm burden , B the prevalence of infection, and C the effective reproductive number , as described in equation 10. Parameter values are set for A. lumbricoides as follows: = 0.81, = 3, = 1 yr, = 0.967 and = 0.95.

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Figure 7.

Effect of regular treatment on mean A. lumbricoides worm burden for different models.

A homogeneous population (left column), B heterogeneous population with uniform transmission dynamics (central column) and C heterogeneous population with greater contribution from children (right column) as in the text. The two rows represent annual and half-yearly treatment respectively. For all runs, basic reproduction number is 3 and worm lifespan is 1 year. Other parameters (as defined for equations 6 and 7): μ2 = 5/yr, k = 0.7, z = 0.93.

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Figure 8.

Effect of regular treatment on mean worm burden of hookworm for different models.

As in Figure 7, the columns are A homogeneous model, B heterogeneous population with uniform transmission dynamics and C heterogeneous population with greater contribution from children, as in the text and different treatment intervals (rows). Simulations for basic reproduction number, = 3, and worm lifespan is 2.5 years. Other parameters as in Figure 7. The two rows represent two-yearly and yearly treatment respectively.

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Figure 9.

Schematic illustration of the impact of school-based deworming on the transmission of parasites.

The number of parasites affected by a school-based deworming programme is never 100%, it is reduced by the efficacy of the drug, the proportion of the population of school age and their intensity of infection, as well as school enrolment and attendance on a deworming day. The impact of such a treatment programme on transmission is less easily quantified and depends on the details of transmission between different age-groups in the population. For further details, see main text.

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