About the Authors

Bronwyn A. Clayton

Affiliation Health and Biosecurity, Australian Animal Health Laboratory, Commonwealth Scientific and Industrial Research Organisation, Geelong, Victoria, Australia

Deborah Middleton

Affiliation Health and Biosecurity, Australian Animal Health Laboratory, Commonwealth Scientific and Industrial Research Organisation, Geelong, Victoria, Australia

Rachel Arkinstall

Affiliation Health and Biosecurity, Australian Animal Health Laboratory, Commonwealth Scientific and Industrial Research Organisation, Geelong, Victoria, Australia

Leah Frazer

Affiliation Health and Biosecurity, Australian Animal Health Laboratory, Commonwealth Scientific and Industrial Research Organisation, Geelong, Victoria, Australia

Lin-Fa Wang

Affiliations Health and Biosecurity, Australian Animal Health Laboratory, Commonwealth Scientific and Industrial Research Organisation, Geelong, Victoria, Australia, Program in Emerging Infectious Disease, Duke–National University of Singapore Graduate Medical School, Singapore

Glenn A. Marsh

Glenn.Marsh@csiro.au

Affiliation Health and Biosecurity, Australian Animal Health Laboratory, Commonwealth Scientific and Industrial Research Organisation, Geelong, Victoria, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BAC DM GAM LFW. Performed the experiments: BAC DM RA LF. Analyzed the data: BAC DM. Wrote the paper: BAC DM.