About the Authors

Stuart D. Dowall

Contributed equally to this work with: Stuart D. Dowall, Victoria A. Graham

stuart.dowall@phe.gov.uk

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Victoria A. Graham

Contributed equally to this work with: Stuart D. Dowall, Victoria A. Graham

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Emma Rayner

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Barry Atkinson

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Graham Hall

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Robert J. Watson

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Andrew Bosworth

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Laura C. Bonney

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Samantha Kitchen

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Roger Hewson

Affiliation National Infection Service, Public Health England, Porton Down, Salisbury, Wiltshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SDD VAG RH. Performed the experiments: SDD VAG ER BA GH RJW AB LCB SK. Analyzed the data: SDD VAG ER BA GH RH. Wrote the paper: SDD VAG ER BA GH RJW RH.